Bret Alexander Beheim

papers I’ve written, or helped write
my full CV as a PDF or as a plaintex YML
also see my google scholar page

- data / code
- citation (bibtex)

Working Papers

Publications in Print


  1. Beheim, B., Bell, A. (2024) Why cultural distance can promote – or impede – group-beneficial outcomes Evolutionary Human Sciences, 6, e14.        

  2. Dalzero, A., Beheim, B., Kaplan, H., Gurven, M., Ross, C.T., Stieglitz, J., Hooper, P.L., Lukas, D. (in press) Cross-cousin marriage among Tsimane forager-horticulturalists during demographic transition and market integration, Evolutionary Human Sciences   


  1. Trumble, B., Negrey, J., Koebele, S.V., Thompson, R.C., Wann, L.S., Allam, A.H., Beheim, B., et al. (2023) Testosterone is positively associated with coronary artery calcium in a low cardiovascular disease risk population Evolution, Medicine, and Public Health, 11 (1), pp.472-484.     

  2. Benjamin C. Trumble, Mia Charifson, Tom Kraft, Angela R. Garcia, Daniel K. Cummings, Paul Hooper, Bret Beheim, et al. (2023) Apolipoprotein-ε4 is associated with higher fecundity in a natural fertility population Science Advances, 9(32), eade9797.     

  3. Hillemann, F., Beheim, B., Ready, E. (2023) Socio-economic predictors of Inuit hunting choices and their implications for climate change adaptation Philosophical Transactions of the Royal Society B, 378(1889), 20220395.        

  4. Stieglitz, J, Buoro, Y, Beheim, B, Trumble, B, Kaplan, H, Gurven, M, et al. (2023). Labour’s pain: strenuous subsistence work, mechanical wear-and-tear and musculoskeletal pain in a non-industrialized population Proceedings of the Royal Society B, 290(1998), 20222497.     

  5. Benjamin C. Trumble, Herman Pontzer, Jonathan Stieglitz, Daniel K. Cummings, Brian Wood, Melissa Emery Thompson, David Raichlen, Bret Beheim, Gandhi Yetish, Hillard Kaplan, Michael Gurven (2023) Energetic costs of testosterone in two physically active foraging populations, American Journal of Human Biology, e23949.       

  6. Thomas S. Kraft, Edmond Seabright, Sarah Alami, Samuel M. Jenness, Paul Hooper, Bret Beheim, et al. (2023) Metapopulation dynamics of infectious disease (SARS-CoV-2) transmission in a small-scale human society PLOS Biology, 21(8).          

  7. Siobhán M. Mattison, Ruizhe Liu, Peter M. Mattison, Bret A. Beheim, Mary K. Shenk, Tami Blumenfield, Chun-Yi Sum, Edmond Seabright, and Sarah Alami (2023) Gender disparities in material and noetic wealth differ by kinship system, Philosophical Transactions B, 378(1883), 20220299.          

  8. Ross, C.T., Hooper, P.L., Smith, J.E., Jaeggi, A.V., Alden Smith, E., Gavrilets, S., Beheim, B. et al. Reproductive inequality in humans and other mammals, Proceedings of the National Academy of Sciences, 120(22), e2220124120.        

  9. Kaplan, H., Hooper, P. L., Gatz, M., Mack, W. J., Law, E. M., Chui, H. C., … & Irimia, A. (2023). Brain volume, energy balance, and cardiovascular health in two nonindustrial South American populations. Proceedings of the National Academy of Sciences, 120(13), e2205448120.     


  1. Fedorova, N., McElreath, R., Beheim, B. The complex life course of mobility: Quantitative description of 300,000 residential moves in 1850-1950 Netherlands. Evolutionary Human Sciences, 4, E39.         

  2. Kraft, T.S., Cummings, D., Venkataraman, V., Alami, S., Beheim, B., Hooper, P. Seabright, E., Trumble, B., Stieglitz, J., Kaplan, H., Endicott, K.L., Endicott, K.M., Gurven, M. (2022) Female cooperative labor networks in hunter-gatherers and horticulturalists Philosophical Transactions of the Royal Society B, 378, 2021043120210431.         


  1. Minocher, R., Atmaca, S., Bavero, C., McElreath, R., & Beheim, B. Estimating the reproducibility of social learning research published between 1955 and 2018. Royal Society Open Science, 8:210450.         

  2. Garcia, A., et al., (2021) APOE4 is associated with elevated blood lipids and lower levels of innate immune biomarkers in a tropical Amerindian subsistence population. eLife, 10:e68231.         

  3. Tran, N.H., Kucharský, S., Waring, T., Atmaca, S., Beheim, B. (2021) Limited scope for group coordination for stylistic variations in kolam art. Frontiers in Psychology, 12:4770.         

  4. Beheim, B., et al. (2021) Treatment of missing data determined conclusions regarding moralizing gods. Nature, 595, pp. E29–E34.         

  5. Jaeggi, A. et al. (2021) Do wealth and inequality associate with health in a small-scale subsistence society? eLife,10:e59437.         

  6. Rowan, C., et al. (2021) Very Low Prevalence and Incidence of Atrial Fibrillation among Bolivian Forager-Farmers. Annals of Global Health, 87(1), p.18.        

  7. Tran, N.H., Waring, T., Atmaca, S., Beheim, B. (2021) Entropy trade-offs in artistic design: A case study of Tamil kolam. Evolutionary Human Sciences, 3(E23).        


  1. Broesch, T., et al. (2020) Navigating cross-cultural research: methodological and ethical considerations Proc. R. Soc. B. 287:20201245.     

  2. Gurven, et al. (2020) Rapidly declining body temperature in a tropical human population Science Advances Vol. 6, no. 44, eabc6599 (credited for data curation)     

  3. Wood, et al. (2020) Step Counts From Satellites: Methods for Integrating Accelerometer and GPS Data for More Accurate Measures of Pedestrian Travel Journal for the Measurement of Physical Behaviour 3(1), pp. 58-66        


  1. Borgerhoff Mulder, et al. (2019) Differences between sons and daughters in the intergenerational transmission of wealth Philosophical Transactions of the Royal Society B 374:20180076        


  1. Ross, et al. (2018) Greater wealth inequality, less polygyny: rethinking the polygyny threshold model Journal of the Royal Society Interface 15:20180035        

  2. Trumble, B.C., Stieglitz, J., Jaeggi, A.V., Beheim, B., Schwartz, M., Seabright, E., Cummings, D., Kaplan, H., and Gurven, M. (2018) Parental hormones are associated with crop loss and family sickness following catastrophic flooding in lowland Bolivia Physiology & Behavior Volume 193, Part A, Pages 101-107.     

  3. Schniter, E., et al. (2018) Information transmission and the oral tradition: Evidence of a late-life service niche for Tsimane Amerindians Evolution and Human Behavior Volume 39, Issue 1, January 2018, Pages 94-105.     


  1. Kaplan, H., et al. (2017) Coronary atherosclerosis in indigenous South American Tsimane: a cross-sectional cohort study The Lancet Volume 389, Issue 10080, pp.1730-1739.     

  2. Gurven, M., Fuerstenberg, E., Trumble, B., Steiglitz, J., Beheim, B., Davis, H., Kaplan, H. (2017) Cognitive performance across the life course of Bolivian forager-farmers with limited schooling Developmental Psychology 53(1), pp. 160–176.     

  3. Blackwell, A.D., Urlacher, S., Beheim, B., von Rueden, C., Jaeggi, A., Stieglitz, J., Trumble, B.C., Gurven, M., Kaplan, H. (2017) Growth references for Tsimane forager-horticulturalists of the Bolivian Amazon American Journal of Physical Anthropology, 162(3), pp.441-461.        

  4. Gurven, Michael, Stieglitz, J., Trumble, B., Blackwell, A.D., Beheim, B., Davis, H., Hooper, P., Kaplan, H. (2017) The Tsimane Health and Life History Project: Integrating anthropology and biomedicine. Evolutionary Anthropology. 26(2) pp.54-73.     

  5. Trumble, B. C., Stieglitz, J., Blackwell, A. D., Allayee, H., Beheim, B., Finch, C. E., Gurven, M., and Kaplan, H. (2017) Apolipoprotein E4 is associated with improved cognitive function in Amazonian forager-horticulturalists with a high parasite burden The FASEB Journal. 31.4, pp. 1508-1515.     


  1. Gurven, M., Costa, M., Eid Rodriguez, D., Stieglitz, J., Beheim, B., Trumble, B., & Kaplan, H. (2016). Health costs of reproduction are minimal despite high fertility, mortality and subsistence lifestyle. Nature Scientific Reports. 6(30056).     

  2. Jaeggi, A.V., Hooper, P.L., Beheim, B.A., Kaplan, H., Gurven, M. (2016) Reciprocal exchange patterned by market forces helps explain cooperation in a small-scale society. Current Biology. 26(16), pp.2180-2187.     

  3. Gurven, M., Trumble, B.C., Stieglitz, J., Yetish, G., Cummings, D., Blackwell, A.D., Beheim, B.A., Kaplan, H., Pontzer, H. (2016) High resting metabolic rate among Amazonian forager-horticulturalists experiencing high pathogen burden. American Journal of Physical Anthropology 161(3), pp.414-425.     

  4. Blackwell, A., Trumble, B.C., Maldonado Suarez, I., Stieglitz, J., Beheim, B.A., Snodgrass, J., Kaplan, H., Gurven, M. (2016) Immune Function in Amazonian Horticulturalists. Annals of Human Biology 43(4), pp. 382-396.     

  5. Mattison, S.M., Smith, E.A., Shenk, M.K., Cochrane, E. (2016) The Evolution of Inequality Evolutionary Anthropology 25.4, pp. 184-199. (acknowledged as data analyst, figure author)        

  6. Mattison, S.M., Beheim, B.A., Chak, B., Buston, P. (2016) Offspring sex preferences among patrilineal and matrilineal Mosuo in Southwest China revealed by differences in parity progression. Royal Society Open Science 3(9) 160526.        


  1. Stieglitz, J., Beheim, B.A., Trumble, B.C, Madiemenos, F.C., Kaplan, H. & Gurven, M. (2015) Low Mineral Density of a Weight-Bearing Bone Among Adult Women in a High Fertility Population. American Journal of Physical Anthropology 156(4), pp.637-648.     

  2. Blackwell, A.D., Tamayo, M.A., Beheim, B.A., Trumble, B.C., Stieglitz, J., Hooper, P.L., Martin, M., Kaplan, H., Gurven, M. (2015) Helminth infection, fecundity, and age of first pregnancy in women. Science 350(6263) pp.970-972.     


  1. Beheim, B. A., Thigpen, C. & McElreath, R. (2014) Strategic Social Learning and the Population Dynamics of Human Behavior. Evolution and Human Behavior 35(5), pp. 351-357        


  1. Beheim, B. A. & Baldini, R. (2012) Evolutionary Decomposition and the Mechanisms of Cultural Change. Cliodynamics 3(2)        


  1. Beheim, B.A. & Bell, A.V. (2011) Inheritance, ecology and the evolution of the canoes of east Oceania. Proc. R. Soc. B 278, pp.3089–3095        

  2. Borgerhoff Mulder, M. & Beheim, B. A. (2011) Understanding the nature of wealth and its effects on human fitness. Phil. Trans. R. Soc. B 366: 344-35        


  1. Borgerhoff Mulder, M., et al. (2009) Intergenerational wealth transmission and the dynamics of inequality in small-scale societies Science Vol. 326, Issue 5953, pp. 682-688 (as data analyst)